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Foundationless frames give bees a blank slate to draw their own comb — but without any guide, they'll build wherever they want, often crosswise between frames. A starter strip runs along the top bar to give bees a straight line to start from, so comb comes out straight and pullable.
Skip the strip and bees will still draw comb — but it often wanders, connecting frames together and making inspections a mess. A thin strip of wax or foundation along the top bar solves that for pennies per frame.